This statistic calculates per tip, the sum of the inverse of all branch lengths of the shortest path between the root and the tip.
References
Williams, P. H., Alonso-Alonso, P., Arbetman, M., Françoso, E., Ghisbain, G., Huang, J., Orr, M. C., Ren, Z.-X., Streinzer, M., T hanoosing, C., Vandame, R., Waite, M., & Brace, S. (2026). Evolutionary Tree for All Bumblebee Species World-Wide Estimated by Combining Information from Fast-Evolving Genes, Slow-Evolving Genes, and Genomic Data (Apidae, Bombus). Insects, 17(6), 540. https://doi.org/10.3390/insects17060540